Mixed stands of Spain
Status:
Stable| Version:v1.0.0(2026-08-28)
General description
- Description: Distance-independent individual-tree growth model for mixed stands in Spain
- Target Species: Pinus sylvestris, Pinus uncinata, Pinus pinea, Pinus halepensis, Pinus nigra, Pinus pinaster, Quercus robur, Quercus petraea, Quercus pyrenaica, Quercus faginea, Quercus ilex, Quercus suber, Fagus sylvatica (codes:
21, 22, 23, 24, 25, 26, 41, 42, 43, 44, 45, 46, 71) - Application Region: Spain
Technical characteristics
- Model Level: Individual tree
- Allows Projection: Yes
- Projection Time Step: 5 years
- Notes: It is always necessary to consider mixed stands of 2 species (see documentation)
To view the step-by-step sequence of initialization, projection, and harvest, see the individual-tree model calculation flow diagram.
Required variables
To run this model in SIMANFOR, the initial inventory must include the following minimum data:
Plot level (Plots sheet)
- Inventory and plot IDs
- species 1 and 2 codes
- dominant height
- Martonne index
Tree level (Trees sheet)
- Inventory, plot and tree IDs
- species
- expansion factor
- dbh
Sample inventories
You can download the following sample inventories in Excel format to test this model:
Modules and equations
Breakdown of simulated biological processes and their methodological references:
- Reference forest growth model: Rodríguez de Prado (2022)
- Growth: Rodríguez De Prado et al. (2022)
- Survival and/or mortality: Rodríguez De Prado et al. (2020)
- Stand Density Index (SDI) reference value: Rodríguez De Prado et al. (2020)
- Height-diameter relationship / height estimation: Vázquez-Veloso et al. (2025), Rodríguez De Prado et al. (2022)
- Tree volume and taper: Badía et al. (2001), Calama & Montero (2006), Fernández-Manso & Sarmiento (2004), Rodríguez et al. (2015), López-Sánchez (2009), Manrique-González et al. (2017), Rodríguez (2009), MITECO (), MITECO (2025)
- Biomass: Diéguez-Aranda et al. (2009), Manrique González et al. (2017), Montero et al. (2005), Ruiz-Peinado et al. (2011), Ruiz-Peinado et al. (2012)
- Carbon content: Castaño-Santamaría & Bravo (2012), Diéguez-Aranda et al. (2009), Herrero de Aza et al. (2011), Montero et al. (2005), Telmo et al. (2010)
- Diversity indexes: Omoro et al. (2010), Simpson (1949), Ulanowicz (2001), Bray & Curtis (1957)
- Structural and size indexes: Varga et al. (2005), Moreno-Fernández & others (2025), Barbeito (2009)
References and documentation
- How to Cite this Model:
SIMANFOR, 2026. Distance-independent individual-tree growth model for mixed stands in Spain (v1.0.0). https://www.simanfor.es
- SIMANFOR Citation:
Bravo, F., Ordóñez, C., Vázquez-Veloso, A., Michalakopoulos, S., 2025. SIMANFOR cloud Decision Support System: Structure, content, and applications. Ecological Modelling 499, 110912. https://doi.org/10.1016/j.ecolmodel.2024.110912
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Technical Documentation: SIMANFOR Models Repository (GitHub)
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Model Bibliography:
- MITECO (). 3er INVENTARIO FORESTAL NACIONAL. DESCRIPCIÓN DE LOS CÓDIGOS DE LA BASE DE DATOS DE SIG. https://www.miteco.gob.es/content/dam/miteco/es/biodiversidad/servicios/banco-datos-naturaleza/documentador_bdsig_ifn3_tcm30-293905.pdf
- MITECO (2025). CUARTO INVENTARIO FORESTAL NACIONAL. Descripción de los códigos de la base de datos Sig. https://www.miteco.gob.es/content/dam/miteco/es/biodiversidad/temas/inventarios-nacionales/documentador_sig_tcm30-536622.pdf
- Badía, M, Rodríguez-Puerta, F, Broto, M (2001). Modelos del perfil del árbol. Aplicación al pino radiata (Pinus radiata D. Don.). https://secforestales.org/publicaciones/index.php/congresos_forestales/article/view/15722/15565
- Barbeito, I. (2009). Natural regeneration dynamics of Pinus sylvestris L. in Central Spain.
- Bray, J. R., Curtis, J. T. (1957). An Ordination of the Upland Forest Communities of Southern Wisconsin. Ecological Monographs.
- Calama, R., Montero, G. (2006). Stand and tree-level variability on stem form and tree volume in Pinus pinea L.: A multilevel random components approach. Forest Systems, 15(1), 24-41. 10.5424/srf/2006151-00951
- Castaño-Santamaría, J., Bravo, F. (2012). Variation in carbon concentration and basic density along stems of sessile oak (Quercus petraea (Matt.) Liebl.) and Pyrenean oak (Quercus pyrenaica Willd.) in the Cantabrian Range (NW Spain). Annals of Forest Science, 69, 663-672. 10.1007/s13595-012-0183-6
- Diéguez-Aranda, U., Álvarez González, J.G., Balboa-Murias, M.A., Barrio-Anta, M., Castedo-Dorado, F., Crecente-Campo, F., González González, J.M., Gorgoso Varela, J.J., López-Sánchez, C.A., Pérez-Cruzado, C., Rodríguez Soalleiro, R., Rojo Alboreca, A., Sánchez Rodríguez, F. (2009). Herramientas selvícolas para la gestión forestal sostenible en Galicia. Consellería do Medio Rural, Xunta de Galicia. https://www.researchgate.net/publication/305443573_Herramientas_selvicolas_para_la_gestion_forestal_sostenible_en_Galicia
- Fernández-Manso, A., Sarmiento, A. (2004). El pino radiata (Pinus radiata). Manual de gestión forestal sostenible. Junta de Castilla y León. https://medioambiente.jcyl.es/web/jcyl/MedioAmbiente/es/Plantilla100Detalle/1284827598811/Publicacion/1284851126859/Redaccion
- Herrero de Aza, C., Turrión, M. B., Pando, V., others (2011). Carbon in heartwood, sapwood and bark along the stem profile in three Mediterranean Pinus species. Annals of Forest Science, 68, 1067. 10.1007/s13595-011-0122-y
- López-Sánchez, C.A. (2009). Estado selvícola y modelos de crecimiento y gestión de plantaciones de Pseudotsuga menziesii (Mirb.) Franco en España. Universidad de Santiago de Compostela,
- Manrique-González, J., Bravo, F., del Peso, C., Herrero, C., Rodríguez, F. (2017). Ecuaciones de perfil para las especies de roble albar (Quercus petraea (Matt.) Liebl.) y rebollo (Quercus pyrenaica Willd) en la comarca de la “Castillería” en el Norte de la provincia de Palencia. http://7cfe.congresoforestal.es/sites/default/files/comunicaciones/776.pdf
- Manrique González, J M, Bravo, F, Del Peso, C, Herrero De Aza, C (2017). Ecuaciones de biomasa para roble albar (Quercus petraea (Matt.) Liebl.) y rebollo (Quercus pyrenaica Willd) en la comarca de la “Castillería” en el Norte de la provincia de Palencia. https://7cfe.congresoforestal.es/sites/default/files/comunicaciones/772.pdf
- Montero, G, Ruiz-Peinado, R, Muñoz (2005). Producción de biomasa y fijación de CO2 por los bosques españoles.. Monografías INIA: Serie Forestal no, 13. https://gregoriomontero.files.wordpress.com/2016/09/2005-01-monografc3ada-forestal-13-m-produccic3b3n-de-biomasa-y-fijacic3b3n-de-co2-por-los-bosques-espac3b1oles.pdf
- Moreno-Fernández, D., others (2025). Structural diversity indices in managed forests.
- Omoro, L. M. A., Starr, M., Pellikka, P. K. E. (2010). Tree biomass and soil carbon stocks in Taita Hills, Kenya. African Journal of Ecology.
- Rodríguez de Prado, Diego (2022). New insights in the modeling and simulation of tree and stand level variables in Mediterranean mixed forests in the present context of climate change. Universidad de Valladolid, https://uvadoc.uva.es/bitstream/handle/10324/55195/TESIS-2015-220907pdf.pdf?sequence=1
- Rodríguez, F. (2009). Cuantificación de productos forestales en la planificación forestal: Análisis de casos con cubiFOR. https://secforestales.org/publicaciones/index.php/congresos_forestales/article/view/17189/17024
- Rodríguez De Prado, Diego, Riofrío, José, Aldea, Jorge, Bravo, Felipe, Herrero De Aza, Celia (2022). Competition and climate influence in the basal area increment models for Mediterranean mixed forests. Forest Ecology and Management, 506, 119955. 10.1016/j.foreco.2021.119955
- Rodríguez De Prado, Diego, San Martín, Roberto, Bravo, Felipe, Herrero De Aza, Celia (2020). Potential climatic influence on maximum stand carrying capacity for 15 Mediterranean coniferous and broadleaf species. Forest Ecology and Management, 460, 117824. 10.1016/j.foreco.2019.117824
- Rodríguez De Prado, Diego, Riofrío, Jose, Aldea, Jorge, McDermott, James, Bravo, Felipe, Herrero De Aza, Celia (2022). Species Mixing Proportion and Aridity Influence in the Height–Diameter Relationship for Different Species Mixtures in Mediterranean Forests. Forests, 13(1), 119. 10.3390/f13010119
- Rodríguez, Francisco, Lizarralde, Iñigo, Bravo, Felipe (2015). Comparison of stem taper equations for eight major tree species in the Spanish Plateau. Forest Systems, 24(3), e034. 10.5424/fs/2015243-06229
- Ruiz-Peinado, R., Montero, G., Del Rio, M. (2012). Biomass models to estimate carbon stocks for hardwood tree species. Forest Systems, 21(1), 42. 10.5424/fs/2112211-02193
- Ruiz-Peinado, Ricardo, Del Rio, Miren, Montero, Gregorio (2011). New models for estimating the carbon sink capacity of Spanish softwood species. Forest Systems, 20(1), 176-188. 10.5424/fs/2011201-11643
- Simpson, E. H. (1949). Measurement of Diversity. Nature, 163, 688.
- Telmo, C., Lousada, J., Moreira, N. (2010). Proximate analysis, backwards stepwise regression between gross calorific value, ultimate and chemical analysis of wood. Bioresource Technology, 101(11), 3808-3815. 10.1016/j.biortech.2010.01.021
- Ulanowicz, R. E. (2001). Information theory in ecology. Computers & Chemistry.
- Varga, P., Chen, H. Y., Klinka, K. (2005). Tree-size diversity in progression of stand development in Douglas-fir forests. Forest Ecology and Management.
- Vázquez-Veloso, Aitor, Yang, Sheng-I, Bullock, Bronson P., Bravo, Felipe (2025). One model to rule them all: A nationwide height–diameter model for 91 Spanish forest species. Forest Ecology and Management, 595, 122981. 10.1016/j.foreco.2025.122981